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What are the Naruto hand seals based on?
There are a number of hand seals in Naruto that are used to perform jutsus. I know that the seals themselves have different zodiac signs. Is there any lore or historical basis for these seals, or was it invented by the creators of Naruto?
The hand seals in Naruto are based in Mudra, which are symbolic/ritual gestures associated to Hinduism and Buddhism. Mudra are used in everyday life, from religious practices to dances, going through martial arts as well. The most wellknown Mudra is probably Añjali Mudrā, which often accompanies the greeting Namaste.
In the French version of the Mudra Wikipedia page, they do indeed link directly to Naruto's hand seals page, unlike what happens in the English page. However, they don't add much info other than the fact that they are called Mudra, and that they're based on the Chinese Zodiac.
From Wikipedia's page on Mudra, on its use on Martial Arts (highlight by me):
One of the more curious things that I encountered in my

The hand seals in Naruto are based in Mudra, which are symbolic/ritual gestures associated to Hinduism and Buddhism. Mudra are used in everyday life, from religious practices to dances, going through martial arts as well. The most wellknown Mudra is probably Añjali Mudrā, which often accompanies the greeting Namaste.
In the French version of the Mudra Wikipedia page, they do indeed link directly to Naruto's hand seals page, unlike what happens in the English page. However, they don't add much info other than the fact that they are called Mudra, and that they're based on the Chinese Zodiac.
From Wikipedia's page on Mudra, on its use on Martial Arts (highlight by me):
One of the more curious things that I encountered in my martial arts training was the use of mudra in combative arts. Mudra (Japanese: in), for those who aren't familiar with them, are these weird hand gestures that are derived from esoteric Buddhism (mikkyo), particularly the Tendai and Shingon sects. These gestu
20171123 05:30:30 
They are based on kujiin. Similar to, but not mudras.
Mudras are learned around peace and happiness while kujiin are developed to handle danger and battle. "Mudras is to qigong" as "kujiin is to budo taijutsu or karate".
20171123 06:10:13